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The in vitro culture of the NCI-H716 human intestinal cell line revealed that nutrients such as palmitic acid, oleic acid and meat hydrolysates can stimulate GLP-1 secretion in a dose-dependent manner (31), however, long-term exposure to long-chain saturated fatty acids such as palmitic acid can lead to ceramide accumulation, caspase-3 activation, and increased DNA fragmentation leading to cell death in GLP-1- producing cells (32)

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